Research articles

30. Geissler, P., Poyarkov, N. A., Grismer, L. L., Nguyen, T. Q., An, H. T., Neang, T., Kupfer, A., Ziegler, T., Böhme, W. & Müller, H. (2015) New Ichthyophis species from Indochina (Gymnophiona, Ichthyophiidae): 1. The unstriped forms with descriptions of three new species and the redescriptions of I. acuminatus Taylor, 1960, I. youngorum Taylor, 1960 and I. laosensis Taylor, 1969. Organisms, Diversity and Evolution 15:143–174.

29. Penske, S., Gvoždík, V., Menegon, M., Loader, S.P. & Müller, H. (2015) Description of the tadpole of Leptopelis cf. grandiceps (Amphibia: Anura: Arthroleptidae) from the Uluguru Mountains, Tanzania. Herpetological Journal 25:61–64.

28. Müller, H. & Larson, J. G. (2014) Description of the tadpole of Ptychadena uzungwensis (Anura: Ptychadenidae) from the Southern Udzungwa Mountains, Tanzania. Journal of Herpetology 48:284–290.

27. Loader, S. P., Ceccarelli, S. F., Menegon, M., Howell, K.M., Kassahun,R., Mengistu, A. A., Saber, S. A., Gebresenbet, F., de Sá, R., Davenport, T. B. R., Larson, J. G., Müller, H., Wilkinson, M. & Gower, D. J. (2014) Persistence and stability of Eastern Afromontane forests: evidence from brevicipitid frogs. Journal of Biogeography 41:1781–1792.

26. Liedtke, H. C., Müller, H., Hafner, J., Nagel, P. & Loader, S. P. (2014) Interspecific patterns for egg and clutch sizes of African Bufonidae (Amphibia: Anura). Zoologischer Anzeiger 253:309–315.

25. San Mauro, D., Gower, D. J., Müller, H., Loader, S. P., Zardoya, R., Nussbaum, R. A. & Wilkinson, M. (2014) Life-history evolution and mitogenomic phylogeny of caecilian amphibians. Molecular Phylogenetics and Evolution 73:177-189.

24. Gower, D. J., Doherty-Bone, T., Loader, S. P., Wilkinson, M., Kouete, M. T., Tapley, B., Orton, F., Daniel, O. Z., Wynne, F., Flach, E., Müller, H., Menegon, M., Stephen, I., Browne, R. K., Fisher, M. C., Cunningham, A. C. & Garner, T. W. J. (2013) Batrachochytrium dendrobatidis infection and lethal chytridiomycosis in caecilian amphibians (Gymnophiona). EcoHealth 10:173-183.

23. Loader, S. P., Menegon, M., Howell, K. H. & Müller, H. (2013) A celebration of the works of John Charles Poynton. African Journal of Herpetology 62:1-4.

22. Wilkinson, M., Gower, D. J., Loader, S. P. & Müller, H. (2013) A visceral synapomorphy of Scolecomorphus Boulenger, 1883 (Amphibia: Gymnophiona: Scolecomorphidae). African Journal of Herpetology 62:21-27.

21. Müller, H., Liedtke, H. C., Menegon, M., Beck, J., Balesteros, L., Nagel, P. & Loader, S. P. (2013) Forests as promoters of terrestrial life-history strategies in East African amphibians. Biology Letters 9:20121146.

20. Kerney, R. R., Blackburn, D. C., Müller, H. & Hanken, J. (2012) Do larval traits re-evolve? Evidence from the embryogenesis of a direct-developing salamander (Plethodon cinereus). Evolution 66:252-262.

19. Loader, S. P., Wilkinson, M., Cotton, J., Measey, G. J., Menegon, M., Howell, K. H., Müller, H. & Gower, D. J. (2011) Molecular phylogenetics of Boulengerula (Amphibia: Gymnophiona: Caeciliidae) and implications for taxonomy, biogeography and conservation. Herpetological Journal 21:5-16.

18. Loader, S. P., Gower, D. J., Müller, H. & Menegon, M. (2010) Two new species of Callulina (Amphibia: Anura: Brevicipitidae) from the Nguru Mountains, Tanzania. Zootaxa 2694:29-42.

17. Müller, H., Wilkinson, M., Loader, S. P., Wirkner, C. S. & Gower, D. J. (2009) Morphology and function of the head in foetal and juvenile Scolecomorphus kirkii (Amphibia: Gymnophiona: Scolecomorphidae). Biological Journal of the Linnean Society 96:491-504.

16. Woltering, J. M., Vonk, F. J., Müller, H., Bardine, N., Tuduce, I. L., de Bakker, M. A., Knöchel, W., Sirbu, I. O., Durston, A. J., Richardson, M. K. (2009) Axial patterning in snakes and caecilians: evidence for an alternative interpretation of the Hox code. Developmental Biology 332:82-89.

15. Sánchez-Villagra, M. R., Müller, H., Scheil, C. A., Scheyer, T. M., Nagashima, H. & Kuratani, S. (2009) Skeletal development in the Chinese soft-shelled turtle Pelodiscus sinensis (Testudines: Trionychidae). Journal of Morphology 270:1381-1399.

14. Kupfer, A., Wilkinson, M., Gower, D. J., Müller, H. & Jehle, R. (2008) Care and parentage in a skin-feeding caecilian amphibian. Journal of Experimental Zoology 309A:460-467.

13. Müller, H., Loader, S. P., Ngalason, W., Howell, K. M. & Gower, D. J. (2007) Reproduction in brevicipitid frogs (Amphibia: Anura: Brevicipitidae) - evidence from Probreviceps m. macrodactylus. Copeia 2007:726-733.

12. Müller, H. (2006) Ontogeny of the skull, lower jaw and hyobranchial skeleton of Hypogeophis rostratus (Amphibia: Gymnophiona: Caeciliidae) revisited. Journal of Morphology 267:968-986.

11. Kupfer, A., Müller, H., Antoniazzi, M. M., Jared, J. C., Greven, H., Nussbaum, R.A. & Wilkinson, M. (2006)A novel form of parental investment by skin feeding in a caecilian amphibian. Nature 440:926-929.

10. Measey, G. J., Mejissa, J. & Müller, H. (2006) Notes on the distribution and abundance of the caecilian Boulengerula uluguruensis in the UluguruMountains, Tanzania. African Journal of Ecology 44:6-13.

9.   Müller, H., Oommen, O. V. & Bartsch, P. (2005) Skeletal development of the direct developing caecilian Gegeneophis ramaswamii (Amphibia: Gymnophiona: Caeciliidae). Zoomorphology 124:171-188.

8.   Müller, H., Measey, G. J., Loader, S. P. & Malonza, P. K. (2005) A new species of Boulengerula Tornier (Amphibia: Gymnophiona: Caeciliidae) from an isolated mountain block of the Taita Hills, Kenya. Zootaxa 1004:37-50.

7.   Müller, H., Measey, G. J. & Malonza, P. K. (2005) Tadpole of Bufo taitanus (Anura: Bufonidae) with notes on its systematic significance and life history. Journal of Herpetology 39:138-141.

6.   Malonza, P. K. & Müller, H. (2004) A rediscovery after two decades: the Changamwe lowland caecilian Boulengerula changamwensis Loveridge, 1932 (Amphibia: Gymnophiona: Caeciliidae). Journal of East African Natural History 93:57-61.

5.   Wilkinson, M., Loader, S. P., Müller, H. & Gower, D. J. (2004) Taxonomic status and phylogenetic relationships of Boulengerula denhardti Nieden, 1912 (Amphibia: Gymnophiona: Caeciliidae). Mitteilungen aus dem Museum für Naturkunde Berlin, Zoologische Reihe 80:41-51.

4.   Kupfer, A. & Müller, H. (2004) On the taxonomy of ichthyophiid caecilians from southern Thailand: A re-evaluation of the holotype of Ichthyophis supachaii Taylor 1960 (Amphibia: Gymnophiona: Ichthyophiidae). Amphibia-Reptilia 25:87-97.

3.   Müller, H. (2003) Zur phylogenetischen Bedeutung der Ontogenese des Schädels von Hypogeophis rostratus und Gegeneophis ramaswamii (Amphibia: Gymnophiona). Sitzungsberichte der Gesellschaft Naturforschender Freunde Berlin (N.F.) 42:57-69.

2.   Wilkinson, M., Müller, H. & Gower, D. J. (2003) On Herpele multiplicata (Amphibia: Gymnophiona: Caeciliidae). African Journal of Herpetology 52:119-122.

1.   Gower, D. J., Kupfer, A., Oommen, O. V., Himstedt, W., Nussbaum, R.A., Loader, S. P., Presswell, B., Müller, H., Krishna, S. B., Boistel, R. & Wilkinson, M. (2002) A molecular phylogeny of ichthyophiid caecilians (Amphibia: Gymnophiona: Ichthyophiidae): Out of India or out of Eurasia? Proceedings of the Royal Society London, Series B 269:1563-1569.

Natural History Notes

4.   Larson, J. G. & Müller, H. (2013) Predation on a Tomopterna luganga (Anura: Pyxicephalidae: Cacosterninae) tadpole by the larva of a water beetle (Dytiscidae: Dytiscinae) in Iringa, southern Tanzania. Herpetology Notes 6:361-362.

3.   Liedtke, H. C. & Müller, H. (2012) Defensive behaviour of Kassina maculata (Anura: Hyperoliidae). Herpetology Notes 5:309-310.

2.   Larson, J. G. & Müller, H. (2011) Pseudotriton ruber. Leucism. Herpetological Review 42:406.

1.   Kupfer, A. & Müller, H. (2001) Ichthyophis kohtaoensis. Maximum length. Herpetological Review 32:249

Books and book chapters

2.   Müller, H. (2007) Developmental morphological diversity in caecilian amphibians: systematic and evolutionary implications. Leiden University Press. Pp. 266.

1.   Müller, H. (2000) Anfang - Franz Eilhard Schulze, der Gründer der Zoologischen Lehrsammlung., p. 183-184, Von der Ästhetik des Lernens II - Die Wandtafeln der Zoologischen Lehrsammlung. p. 192-193, plus 30 additional object descriptions pp. 152, 185-191, 195-196 in: Bredekamp, H, Brüning, J & Weber, C. (eds.) Theater der Natur und Kunst - Theatrum naturae et artis: Wunderkammern des Wissens. Katalog. Henschel, Berlin. Pp. 311. [Catalogue to: "Theatre of Nature and Art - Treasure-trove of Knowledge" An exhibition organised by the Humboldt-University Berlin under the patronage of the President of the Federal Republic of Germany, held at the Martin Gropius Bau, Berlin from December 10, 2000 to March 4, 2001.]

Popular articles

4.   Measey, G. J., Müller, H. & Loader, S. P. (2013) African Journal of Herpetology celebrates John Poynton. Froglog 107:48.

3.   Loader, S. P., Menegon, M., Müller, H., Gower, D. J., Wilkinson, M., Howell, K., & Orton, F. (2009) Remarkable amphibian diversity in the South Nguru Mountains, Tanzania. Froglog 91:8-10.

2.   Kupfer, A., Müller, H., Measey, G. J., & Malonza, P. K. (2005) Auf Blindwühlensafari in Ostafrika. Amphibia 4:8-11.

1.   Malonza, P. K. & Müller, H. (2005) For so long in the dark. SWARA 28:34-35.

Book reviews

4.   Müller, H. (2012) Amphibians of Malawi, by V. Mercurio. Herpetological Review 43:160-161.

3.   Müller, H. (2011) The Amphibians and Reptiles of Ethiopia and Eritrea, by M. Largen & S. Spawls. Herpetological Review 42:309-311.

2.   Müller, H. (2007) Reproductive Biology and Phylogeny of the Gymnophiona, by Jean-Marie Exbrayat (ed). Acta Zoologica 89:89-91.

1.   Müller, H. (2006) Amphibians of East Africa, by A. Channing & K. M. Howell. Mitteilungen aus dem Museum für Naturkunde Berlin, Zoologische Reihe 82:307-308.

Species conservation assessments (available at

2.   Measey, J., Malonza, P. & Müller, H. (2006) Boulengerula niedeni. In: IUCN 2006. 2006 IUCN Red List of Threatened Species.

1.   van Dijk, P.P., Chuaynkern, Y., Kupfer, A. & Müller, H. (2004) Ichthyophis supachaii. In: IUCN 2006. 2006 IUCN Red List of Threatened Species.